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.1996.,
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.244-239
:3
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.1990.,.,,.,-,.,
.178-169
:31..
.2
.34-31:'.19.1983.,.,
.3
.1996.,.,,.,,.,
.187-177
:3..
.4

5.
Anderson,D.C.,Harper,K.T.andHolmgren,R.C.1982.Factors
influencingdevelopmentofcryptogamicsoilcrustsinUtahdesert.J.
RangeManage.35:180-185.
6.
Bar-Or,Y.andDanin,A.1989.Mechanismsofcyanobacterialcrust
formationandsoilstabilizationinthenorthwesternNegev,Israel.
SocietyofMicrobiology,Haifa,13pp.
7.
Bar-Or,Y.andShilo,M.1988.Theroleofcell-boundflocculantsin
coflocculationofbenthiccyanobacteriawithclayparticls.FEMS
MicrobiologicalEcology.53:169-174.
8.
Belnap,J.1993.Recoveryratesofcryptogamiccrusts:inoculant
useandassessmentmethods.GreatBasinNaturalist.53:89-95.
9.
Boeken,B.andShachak,M.1995.Changesindesertplant
communitiesinhuman-madepatchesandtheirimplicationsfor
managementofdesertifiedlandscapes.Ecologicalapplication.4:
702-716.
10
.
Booth,W.E.1941.Algaeaspioneerinplantsuccessionandtheir
importanceinerosioncontrol.Ecology.22:38-46.
11
.
Brock,T.D.1975.EffectofwaterpotentialonaMicrocoleus
(Cyanophyceae)fromdesertcrust.J.ofPhycology.11:316-320.
12
.
Buzer,J.S.,Dohmeier,R.A.andTroitduD.R.1985.Thesurvival
ofalgaeindrysoilsexposedtohightemperaturesforextendedtime
periods.Phycologia.24:249-251.
13
.
S.,andGolubic,S.1989.Desertcrust-Campbell,S.E.,Seeler,J
formationandsoilstabilization.AridSoilResearchand
Rehabilitation.3:217-228.
14
.
Danin,A.1996.Plantsofdesertdunes.Springer-Verlag,Berlin.p.
177.
15
.
Danin,A.andYaalonD.H.1982.Siltplusclaysedimentationand
decalcificationduringplantsuccessioninsandsofMediterranean
coastalareaofIsrael.IsraelJ.ofEarthScience.31:101-109.
16
.
Eldridge,D.J.1994.Microbioticsoilcrusts:Areviewoftheirrolesin
soilandecologicalprocessesintherangelandsofAusteralia.
AusteralianJ.ofSoilResearch32:389-415.
17
.
Friedman,E.I.andGalun,M.1974.Desertalgae,lichensandfungi.
In:Brown,
G.W.(Ed.),DesertBiology.AcademiePress,London.Vol.2,pp.
165-212.
18
.
Gutterman,Y.,Golan,T.andGarsani,M.1990.Porcupinediggings
asauniqueecologicalsysteminadesertenvironment.Oecologia
85:122-127.
19
.
Harper,K.T.,andpendleton,R.S.1993.Cyanobacteriaand
cyanobacteriallichens:cantheyenhanceavailabilityofessential
mineralsforhigherplants?GreatBasinNaturalist.53:59-72.
20
.
Jefferies,D.L.,andKloptek,J.M.,Link,S.O.,andBolton,H.Jr.
1992.AcetylenReductionbycryptogamiccrustsfromablackbrush
communityasrelatedtoresaturationanddehydration.SoilBiology
Biochemistry24:1101-1105.
21
.
JohansenJ.R.1986.Importanceofcryptogamicsoilcruststoarid
rangelands:implicationsforshortdurationgrazing.InTiedeman,J.
A.(Ed.),ShortDurationGrazing.WashingtonStateUniv.Pullman,
Washington.pp.127-136.
22
.
Johansen,J.R.1993.Minireview:Cryptogamiccrustsofsemiarid
andaridlandsofnorthAmerica,J.ofPhycology29:140-147.
23
.
Johansen,R.J.,andSt.Clair,L.L.1986.Cryptogamicsoilcrusts:
recoveryfromgrazingnearcampfloydstatepark,Utah,U.S.A.
GreatBasinNaturalist,46:632-640.
24
.
Kleiner,E.F.andHarper,K.T.1977.Occurrenceoffourmajor
perennialgrassesinrelationtoedaphicfactorsinapristine
community:J.RangeManagement30:286-289.
25
.
Kutiel,P.,Danin,A.andOrshan,G.1979/1980.Vegetationofthe
sandysoilsnearCaesarea,Israel.I.Plantsuccession.Israel.J.
Botany28:20-35.
26
.
Lange,O.L.Kidron,G.J.,Budel,B.,Mayer,A.,Killian,E.and
Abeliovitch,A.1992.Taxonomiccompositionandphotosynthetic
characteristicsofthe'biologicalsoilcrusts'coveringsanddunesin
thewesternNegevDesert.FunctionalEcology6:519-527.
27
.
Levy,Y.andSteinberger,Y.1986.Adaptationofdesertsoil
microalgaetovaryinglightintensities.CompativePhysiologyand
Ecology11:90-94.
28
.
Marble,J.R.,Harper,K.T.1989.Effectoftimingofgrazingon
soil-surfacecryptogamiccommunitiesinaGreatBasinlowshrub
desert:Apreliminaryreport.GreatBasinNaturalist.49:104-107.
29
.
Metting,B.1991.Biologicalsurfacefeaturesofsemiaridlandand
deserts.In:Skujins,J.(Ed.),SemiaridlandsanddesertsSoil
resourceandreclamation.MarcelDeckerInc.,NewYork.pp.
257-293.
30
.
Metting,B.andRayburn,W.1979.Theeffectsofthe
pre-emergenceherbicidedi-allateandthepost-emergence
herbicideMCPAongrowthofsomesoilalgae.Phycologia.18:
269-272.
31
.
Rubio,H.O.,Wood,M.K.,Cardenas,M.andBuchanan,B.A.
1992.Theeffectofpolyacrylamideongrassemergencein
southcentralNewMexico.J.RangeManagement.45:296-300.
32
.
Rychert,R.C.andSkujins,J.1974.NitrogenFixationbyblue-green
algae-lichencrustsintheGreatBasinDesert.SoilScienceSociety
ofamericaProceedings38:768-771.
33
.
Schofield,W.B.1985.Intoductiontobryology.Macmillian,New
York.
34
.
Shachak,M.1980.Energyallocationandlifehistorystrategyofthe
desertisopod,Hemilepistusreaumuri.Oecologia(Berl.).45:
404-413.
35
.
Shachak,M.andSteinberger,Y.1980.Analgae-desertsnailfood
chain:Energyflowandsoilturnover.Oecologia(Berl.).46:402-411.
36
.
St.Clair,L.L.,Johansen,J.R.,andWebb,B.L.1986.Rapid
stabilizationoffire-disturbedsitesusingasoilcrustslurry:
inoculationstudies.Reclam.Revege.Research4:261-269.
37
.
West,N.E.1990.Structureandfunctionofmicrophyticsoilcrustsin
wildlandecosysteofaridtosemi-aridregions.AdvancesinEcological
Research20:179-223.
38
.
Yair,A.1990.Runoffgenerationinasandyarea-theNizzana
sands,WesternNegev,Israel.EarthSurfaceProceedings15:
597-609.
39
.
Yair,A.andShachak,M.1987.Studiesinwatershedecologyonan
aridarea.In:M.O.WurteleandBerkofsky,L.(Eds.).Progressin
DesertResearch.KrowmanandLittlefieldPub.pp.146-193.
40
.
Zaady,E.,Groffman,P.andShachak,M.1996.Releaseand
consumptionofsnailfecesinNegevdesertsoils.Biology&Fertility
ofSoils.23:399-404.
41
.
Zaady,E.,Groffman,P.andShachak,M.1998.Nitrogenfixationin
macro-andmicrophyticpatchesintheNegevdesert.SoilBiology&
Biochemistry30:449-454.
42
.
Zaady,E.,Okon,Y.andPerevolotsky,A.1994.Growthresponseof
MediterraneanherbaceousswardstoinoculationwithAzospirillum
brasilense.J.ofRangeManagement47:12-15.
43
.
Zaady,E.andShachak,M.1994.Microphyticsoilcrustand
ecosystemleakageintheNegevDesert.AmericanJ.ofBotany
87:109.
44
.
Zaady,E.,Gutterman,Y.andBoeken,B.1997.Thegermination
effectsofcyanobacterialsoilcrustonmucilaginousseedsofthree
desertsplants:Plantagocoronopus,Reboudiapinnataand
Carrichteraannua.PlantandSoil.190:247-252.





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